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The soldier caste has anatomical and behavioural specialisations, and their only real aim is to defend the colony. Many soldiers have big heads with exceptionally altered strong jaws therefore enlarged they cannot feed themselves. Instead, such as juveniles, they're fed by employees.5556 Fontanelles, easy holes in the eyebrow which exude defensive secretionsare a feature of the family Rhinotermitidae.57 Many species are readily identified using the characteristics of the soldiers' bigger and darker head and massive mandibles.53 Among certain termites, soldiers can utilize their globular (phragmotic) heads to block their narrow tunnels.58 Different sorts of soldiers include minor and major soldiers, and nasutes, which have a horn-like nozzle frontal projection (a nasus).53 These unique soldiers can spray noxious, sticky secretions containing diterpenes in their enemies.59 Nitrogen fixation has an important role in nasute nutrition.60.

The reproductive caste of a mature colony includes a fertile female and male, known as the queen and king.61 The queen of this colony is responsible for egg production for its colony. Unlike in ants, the king mates with her life.62 In some species, the abdomen of this queen swells up radically to increase fecundity, a characteristic known as physogastrism.61 Depending on the species, the queen starts producing reproductive winged alates at a certain time of year, and huge swarms emerge from the colony when nuptial flight begins.

A young termite nymph. Nymphs first moult into employees, but others might further moult to become soldiers or alates.

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Termites are often compared with the social Hymenoptera (ants and various species of bees and wasps), but their differing evolutionary origins result in major differences in life span. In the eusocial Hymenoptera, the workers are entirely female. Men (drones) are haploid and develop from unfertilised eggs, while females (both workers and the queen) are diploid and grow from fertilised eggs.

Depending on species, both male and female employees may have different roles in a termite colony.63.

The entire life span of a termite begins with an egg, but is different from that of a bee or ant in that it goes through a developmental check that process called incomplete metamorphosis, with egg, nymph and adult stages.64 Nymphs resemble little adults, and go through a series of moults as they grow.

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The development of nymphs into adults can take months; the time period depends on food availability, temperature, and the general population of this colony. Since nymphs are unable to feed themselves, workers must feed them, but workers also take part in the social life of the colony and have certain other tasks to achieve such as foraging, building or maintaining the nest or tending to the queen.5367 Pheromones govern the caste system in termite colonies, preventing all but a very few of the termites from becoming fertile queens.68.

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Queens of the eusocial termite Reticulitermes speratus are effective at a long lifespan without sacrificing fecundity. These long-lived queens have a significantly lower level of oxidative damage, including oxidative DNA damage, than employees, soldiers and nymphs.69 The lower degrees of damage appear to be due to increased catalase, an enzyme that protects against oxidative stress.69.

Termite alates only depart the colony when a nuptial flight occurs. Alate males and females pair up together and then land in search of a suitable spot for a colony.70 A termite king and queen do not mate until they find such a spot. When they dothey excavate a room big enough for both, close up the entrance and move to mate.70 After mating, the pair never go outdoors and spend the rest of their lives in the nest.

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For example, alates in certain species emerge during the day in summer while some emerge during winter.71 The nuptial flight might also begin at dusk, when the alates swarm around regions with lots of lights. The time when nuptial flight begins depends on the environmental conditions, the time of day, moisture, wind speed and precipitation.71 The number of termites in a colony also fluctuates, with the larger species typically having 1001,000 individuals.

The queen only lays 1020 eggs in the very early stages of the colony, but lays as many as 1,000 per day when the colony is a few years old.53 At adulthood, a primary queen has a great capacity to lay eggs. In certain species, the adult queen includes a greatly distended abdomen and may create 40,000 eggs per day.72 Both adult ovaries may have some 2,000 ovarioles each.73 The abdomen increases the queen's body length to several times more than prior to mating and reduces her ability to move freely; attendant workers provide assistance.

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